Showing posts with label arilbred. Show all posts
Showing posts with label arilbred. Show all posts

Monday, April 3, 2023

The Aril Society Checklist

 by Tom Waters

When I'm not writing blog posts, one of my many other iris-related activities is serving as checklist editor for the Aril Society International. I've just completed the 2023 update of the checklist, which is available on the ASI website. It occurred to me that there are probably many people who are not aware that the ASI maintains a checklist of aril and arilbred irises, nor aware of what it offers beyond other references. So I thought I would use this blog post to have a look at the checklist and why it is important.

When the Aril Society International was formed in the 1950s, there were no standardized definitions for the various types of aril and arilbred irises. Irises with oncocyclus or Regelia ancestry were registered with the American Iris Society under a bewildering range of classification codes. One of the first tasks of the new society was to sort through all the various registrations and establish specific categories for them. A persistent problem in the early years was that irises of 1/8, 1/16, or even less aril ancestry were being given awards as arilbreds despite being indistinguishable from TBs. The society addressed this by requiring irises to be 1/4 aril or more to be considered as arilbreds. Subsequently, there were two separate award systems created: one for arilbreds of 1/4 to 1/2 aril content, and another for irises of 1/2 aril content or more.

'Loudmouth' (Rich, 1970) won the
C. G. White Award as an OB (1/2 aril)
but is now an OB- (less than 1/2) 
because of a change in definition.

The checklist was critical for establishing which irises met the various award criteria. The editor would take the registration data, do pedigree research, consider an iris's appearance, breeding behavior, and any chromosome counts that had been done, and then assign it to the appropriate category. In days before the internet, the ASI checklist was the only source of this classification information, because it was impractical to reprint the American Iris Society checklists to give updated classifications for arilbreds. The classifications and definitions used by the Aril Society have changed several times over the years, and the checklist is the only authoritative reference on how definition changes have affected the classification of each arilbred iris. Applying the current definitions, which involve estimating the chromosome makeup of each cultivar, is not a trivial matter in some cases.

Here are some of the things you will find in the ASI checklist that may not be presented accurately (or at all) in other sources:

Classification Information. Each iris listed, even historic irises registered long ago under other definitions, are assigned to the current ASI classification, to the extent possible given the existing information. This is of particular value to show classification committees in regions where arils and arilbreds are likely to be exhibited. It is especially important for irises that were once considered arilbreds, but no longer meet the definitions of that class.

Height Categories. In 2018, the ASI established definitions for two height categories for smaller arilbreds: arilbred dwarf (ABD) and arilbred median (ABM). These are included in the current edition of the AIS Judges' Handbook. The checklist gives the correct height category for each arilbred listed.

Chromosome Configuration. Aril and arilbred irises may include chromosome sets from oncocyclus, Regelia, tall bearded, and dwarf bearded species. Which chromosome sets are present in a given iris is of considerable interest to hybridizers, as it affects fertility and helps the hybridizer properly classify any resulting seedlings. The checklist provides the most likely chromosome configuration for most irises listed, along with an actual chromosome count where one has been reported.

Fertility Observations. When an iris is known to have produced offspring as a seed parent or pollen parent, this is noted in the checklist.

Is this 'Bronze Beauty',
'Hoogiana Bronze Beauty',
Iris hoogiana 'Bronze Beauty',
'Bronze Beauty Van Tubergen',
or 'Antiope'? Answer: yes.
Irises Not Eligible for Awards as Arilbreds. An appendix to the checklist provides a thorough listing
of irises that do not qualify as arilbreds for various reasons: never having been registered, having been registered in a non-arilbred class, or having been disqualified as definitions became more strict.

Nonregistered Aril Hybrids. A substantial number of aril hybrids have been introduced, particularly through the firm of Van Tubergen in the Netherlands, that were never registered with the American Iris Society. Some were registered with the Dutch registration authority for bulb irises, while others were simply not registered at all. The checklist includes these in an appendix, along with photographs and other information.

Even today, where so much information seems just clicks away on the internet, the ASI checklist fills an important role in providing authoritative information that is not reliably available elsewhere. It is an important resource for judges, show officials, and hybridizers. Anyone with an interest in aril and arilbred irises should become familiar with what the checklist has to offer.




Monday, May 31, 2021

Arilpums Are Here!

by Tom Waters

When I last wrote about my progress with hybridizing arilpums in September, I was eagerly anticipating this spring's bloom, with many new seedlings expected to bloom on established plants. I was not disappointed!

Aladdin's Gem

First, a little recap. Arilpums are hybrids between pure arils and the dwarf bearded species Iris pumila. When a tetraploid aril is used as the aril parent, theory predicts that the arilpum seedlings will be fully fertile amphidiploids. Historically, most arilbred dwarfs and medians have been produced by using arilbreds, diploid arils, and SDBs in various combinations, and the result has been infertile offspring, making it impossible to linebreed the smaller arilbreds to refine them and extend their potential. Arilpums are the way out of that difficulty. However, only a few have been registered and introduced: 'Barbarella' (Mathes, 1991), 'Aladdin's Gem' (Thoolen, 2002), and 'Topaz Talisman' (Jensen, 2015). Although producing fertile pollen, arilpums so far have shown little or no pod fertility, a disappointing setback.

Last spring, I had some luck with the two arilpum seedlings I bloomed, Q025-01, 'Merlin's Magic' X Armenian pumila seedling, and Q053-01, 'Tadzhiki Bandit' X 'Sudden Butterflies'. Both produced seeds for me, as did a seedling of George Hildenbrandt's, GH-11-4-2, 'Dunshanbe' X 'Hidden Dragon'. So it seemed there was hope after all for creating a line of fertile arilpums!

This spring reinforced my optimisim, and also gave a preview of the great variety of color, pattern, and form that is possible with arilpums.

Q068-14


Q068-04, with "monster beards"











Earliest to bloom were seedlings from cross Q068, 'Dunshanbe' X 'Wild Whispers'. I had over a dozen of these, most dark violet selfs with some aril look but without much distinctiveness. Two stood out, however. Q068-04 has massive caterpillar beards. Not exactly beautiful, but certainly memorable. Q068-09 is the best of the lot, with a clean oncocyclus shape and elegant look. 

Q068-09, the standout

The second cross to bloom was really exciting: Q066, 'Tadzhiki Bandit' X Caucasus pumila seedling. These were much smaller and more varied than the 'Dunshanbe' seedlings, showing many exciting colors and patterns. I've kept almost everything from this cross for further evaluation.

Q066-06

Q066-02

There was another from cross Q053, looking almost the same as its sibling that bloomed last year. I also had a few seedlings from cross Q070, 'Vera' X 'Royal Wonder', one of which was deemed worthy of a closer look.

Q070-01

I made a number of crosses with this year's arilpums, but was not very systematic or thorough about it. Seeds have not been harvested yet, so it is premature to say much about their pod fertility. But there are pods, even some bee pods, so I am optimistic. 

It's been about ten years work to get to this point, but at last I have a small cadre of arilpum seedlings to work with, and the future looks very promising.




Monday, September 14, 2020

Progress with Arilpums

by Tom Waters

When I wrote my previous blog post on arilpums early in 2019, I had to focus mostly on history and theory, as I did not yet have much to show from my own seedlings. That has now changed, and the current state of this project is even better than I had dared hope.

As explained in the previous post, arilpums are arilbreds dwarfs that come from crossing pure arils with the dwarf bearded species Iris pumila. If the aril parent is a tetraploid, theory says the resulting arilpums should be fertile amphidiploids, a sort of arilbred analog of the SDBs. They thus have the potential of ushering in a whole new "fertile family" of miniature arilbreds.

Waters Q025-01

This spring, I bloomed two arilpum seedlings, both prolific two-year-old clumps covered in bloom. The first, Q025-01, from 'Merlin's Magic' (a selection of Iris stolonifera) x P002-05, a pumila seedling of Armenian origin, has a dainty Regelia look and some nice veining. The second seedling, Q053-01, from 'Tadzhiki Bandit' x 'Sudden Butterflies' is larger and huskier, with flowers reminiscent of tall arilbreds with Regelia ancestry, such as 'Stars Over Chicago' or 'Saffron Charm'.

I also had good bloom this year on an arilpum seedling from

Waters Q053-01

George Hildenbrandt, GH-11-4-2, from 'Dunshanbe' x 'Hidden Dragon'. This is a charming dark arilpum with nice aril-like flower form.

Hildenbrandt GH-11-4-2

The remarkable thing is that all three of these seedling produced seed for me this year when crossed with other arilpums. In my previous post, I mentioned that earlier hybridizers working with arilpums had found them to be infertile as pod parents. That is clearly (and thankfully!) not a general rule! Here are my successful arilpum x arilpum crosses this year:

Q053-01 x Aladdin's Gem gave 38 seeds
Q025-01 x Q053-01 gave 9 seeds
GH-11-4-2 x Q053-01 gave 23 seeds
GH-11-4-2 x Aladdin's Gem gave 3 seeds

These are not huge seed yields, but they are certainly adequate for continued breeding. I am quite optimistic that a number of the second-generation seedlings from these crosses will be fully fertile, and it will be possible to line breed arilpums without having to constantly go back to the difficult aril x pumila initial cross. This will also open up arilpum breeding for hybridizers who live in areas where either arils or pumila are difficult to grow.

Next spring should offer even more excitement, as I have more than 30 arilpum seedlings from various crosses that will be blooming as two-year-old plants. I got a sneak preview of one this spring,  Q066-08, from 'Tadzhiki Bandit' x P018-02, a pumila seedling originating from the Caucasus mountains. It looks for all the world like a miniature rendition of its aril pod parent, giving a rather more "onco" impression than others mentioned in this post. It has not yet been tested for fertility.

Waters Q066-08

All this has been an exciting vindication of a breeding project motivated mostly by theoretic possibilities rather than concrete experience. I'm now confident that arilpums have a bright future ahead.


Monday, March 23, 2020

Irises in Containers

by Tom Waters

Irises are not usually thought of as container plants, but they can grow quite well that way, and there are a number of advantages to doing so.

Two Iris pumila cultivars,
'Wild Whispers' (Coleman, 2012) and
'Royal Wonder' (Coleman, 2013),
growing happily in a large container
The irises I choose to grow in containers are mostly dwarf bearded irises and the smaller arils and arilbreds. Tall bearded irises look out of place in even the largest containers, and a light container soil mix may not give them the support they need when top-heavy with bloom. The smaller irises, however, are naturals for container culture. They bring the plants closer to eye level for viewing, and allow them to be moved from place to place for best effect. If attractive containers, like oak barrels or terracotta pots are used, the effect can be quite lovely and dramatic.

There are additional advantages to container culture. I tend to put rare or choice plants into containers when I first acquire them, as it makes them much easier to weed and care for. A small iris that might get lost in an overgrown summer garden and succumb to neglect, is kept safe in a container where it can get the attention it needs.

Two forms of the exotic oncocyclus species Iris paradoxa
in a large container with a gritty soil mix
Arils are another good candidate for container culture in climates where summers are too wet to grow them successfully in the ground. The container can be stored in a warm dry place through the irises' summer dormancy period. It is also possible to provide a coarse, well-draining soil mix that would be difficult to maintain in the open garden.

As a hybridizer working with dwarfs and other small irises, I also appreciate that containers make the blooms more accessible. It's much nicer to pull a chair up to a container than to crawl around on the ground to harvest pollen or make a cross.

Most of my containers are inexpensive plastic models, in the largest size possible (two to three feet in height and diameter). Even the smallest irises enjoy a wide and deep root run. I fill them with various soil mixes, depending on what I have at hand, but I usually use a mix of my garden soil (a somewhat sandy silt) and commercial potting soil, sometimes with addition of compost, coarse sand, or even small gravel. The irises do not seem too picky about the exact composition of the potting mix. I like to use a top dressing of gravel as a mulch. It also looks nice, especially if a few rocks are positioned on top to make a miniature landscape.

Even in a container with enriched soil, irises will not go forever without dividing them and refreshing the soil mix every few years. Keep an eye out for overcrowding or declining vigor. Also, it is important to keep to a regular watering schedule; how frequently you water will depend on your climate, but the only time I have lost an otherwise healthy plant in a container was when I accidentally let it get bone-dry in the summer. Containers are less forgiving in this way than garden soil.

If you've never grown irises in containers, give it a try this year! You may find it offers both esthetic and practical rewards.
Iris reichenbachii blooming profusely in a container


Monday, August 19, 2019

2019 CG White Medal

The American Iris Society
Announces
THE CLARENCE G. WHITE MEDAL
'Dubai'

'Dubai'--image by Paul Black

'Dubai' ( Thomas Johnson, R. 2013) AB, OGB. Seedling TC229A. AB (OGB+), 33 (84 cm), Early to midseason bloom. Standards and style arms light lilac-purple; Falls light strawberry-rose, large red-black signal; beards yellow-orange; slight fragrance. 'Lancer' x 'Energizer'. Mid-America 2013. Honorable Mention 2015. Award of Merit 2017.

Since 1993, the Clarence G. White memorial medal has been awarded to the best arilbred iris with 1/2 or more aril ancestry. When Clarence G. White began his work with aril irises, little was known about the complexity of iris genetics. White assembled the largest collection of aril irises in the world, and conducted thousands of breeding experiments to obtain viable, fertile seedlings. One of his goals was to develop strong, pure Oncocylus hybrids.

The World of Irises blog will be posting once a day all of the medal winners. The entire list of winners can be found at the AIS website, the AIS Encyclopedia and later in the AIS Bulletin, IRISES.

Monday, April 8, 2019

Arilpums: Fertile Arilbred Dwarfs!

by Tom Waters

'Aladdin's Gem' (Thoolen, 2002)
OGB/ABD

Most arilbred medians are produced by crossing standard dwarfs (SDBs) with fertile halfbreds (OGB). The results of such crosses tend to be pleasing garden subjects, occupying about the same niche as intermediates (IBs), but with the added interest of arilbred color patterns. Unfortunately, arilbred medians produced this way are usually infertile, or have very limited fertility at best. Also, being only ¼ aril (OGB-), they don’t always show enough pronounced aril characteristics to really stand out as arilbreds.

For those of us who enjoy the smaller arilbreds, the holy grail would be diminutive fertile arilbreds that are half aril in content. We might imagine miniaturized versions of the taller arilbreds that have come from breeding arils with tall bearded (TBs). One approach to realizing this dream is through the creation of “arilpums”: hybrids between arils and the tiny dwarf bearded species Iris pumila. This type of cross produces true arilbred dwarfs, around 10-12 inches (25-30 cm) in height, fertile, and half aril. Additionally, Iris pumila is expected to confer winter hardiness on its seedlings.

The potential of such breeding was first appreciated in the mid 20th century, when iris breeders became aware of the cytogenetics of irises, and the possibility of creating “fertile families” of irises from wide crosses, if the parents are chosen to have the right chromosome configurations to produce fertile offspring. Both the SDBs (from TBs crossed with Iris pumila) and the fertile arilbreds (from TBs crossed with arils) are examples of such fertile families, and show how varied and successful they can be.

The most direct way to build a fertile family of arilpums is to cross Iris pumila with tetraploid arils. In nature, the only tetraploid arils are the Regelia species Iris hoogiana and Iris stolonifera. Indeed, the pioneering hybridizer Paul Cook (who also introduced the first SDBs from TB x pumila crosses) produced two hoogiana/pumila hybrids in the 1950s: ‘Hoogpum Blue’ (Cook, 1956) and ‘Hoogpum Purple’ (Cook, 1956). Both were fertile, but Iris hoogiana doesn’t have many characteristics that most of us think of as epitomizing the “aril look”, and these little offspring were not very distinguishable from MDBs, and in fact were registered as such.

The first approach to arilpums with oncocyclus ancestry came from Peter Werckmeister, the great expositor of cytogenetic theory to English-speaking irisarians. In the 1970s, he produced “Miltonia” (not registered) from a lucky cross, in which the aril parent (an oncogelia from ‘Teucros’ x Iris susiana), although a diploid, acted as a tetraploid by producing an unreduced gamete. Werckmeister reported eventually creating a whole array of fertile arilpums from “Miltonia”. Alas, they were not widely distributed and are now lost to us.

Thoolen arilpum seedling AP9007-1
Interest in arilpums underwent something of a revival in the 1990s, after Samuel Norris and John Holden created a line of tetraploid oncogelias from using colchicine treatment. Crossing these with Iris pumila produced arilpums with much more “onco look” than had been possible previously. The first of these to be introduced was ‘Barbarella’ (Mathes, 1991), followed by ‘Aladdin’s Gem’ (Thoolen, 2002). Both Harald Mathes and Francesca Thoolen raised a number of different arilpum seedlings, but these were the only two registered and introduced into commerce. Both ‘Barbarella’ and ‘Aladdin’s Gem’ produce fertile pollen, but are unwilling pod parents. Mathes and Thoolen noted similar difficulties with their other arilpums, and their breeding programs were eventually abandoned.

The lack of pod fertility was an unfortunate setback, because the simplest way to build up this family would be to cross the existing arilpums among themselves, rather than repeating the initial aril x pumila crosses.

'Topaz Talisman' (Jensen, 2015)
RB/ABD
The most recent arilpum is the lovely stolonifera/pumila hybrid ‘Topaz Talisman’ (Jensen, 2015). It has not yet been tested extensively for pod fertility.

Why has this family not made more progress? It has been more half a century since its potential was first understood, theoretically. I do not believe the pod fertility issue is insurmountable, although it is a setback and perhaps kept the family from taking off in the 1990s, when interest was greatest. Since then, the number of hybridizers working with arilbreds has declined, and not all are interested in such an experimental project. The parent plants, tetraploid arils and Iris pumila, are not easy to grow in all locations: the arils need a very dry summer, and pumila needs a cold winter.

Another obstacle is that the initial crosses are difficult, not many viable seeds are produced, germination is poor, and so a lot of persistence is needed to bring arilpum seedlings to bloom. Since I began working in this area six years ago, I have made 365 pollinations, resulting in 525 seeds, and at long last four seedlings which I hope to see bloom this year or next.

I would love to see others get involved; the prospects for eventual success obviously depend on a number of people working with determination over the course of a number of years. Here is the recipe:
  1. Acquire tetraploid arils, such as Iris stolonifera, ‘Tadzhiki Bandit’, ‘Tadzhiki Eclipse’, ‘Werckmeister’s Beauty’, ‘Balalaika Music’, and ‘Dunshanbe’. Keep in mind that you must use tetraploid arils – crossing Iris pumila with random arils and arilbreds offers no special hope of fertility in the seedlings.
  2. Acquire Iris pumila, either from seed exchanges, or named cultivars such as ‘Royal Wonder’, ‘Wild Whispers’, ‘Little Drummer Boy’, ‘Hobbit’, and others.
  3. Make crosses between these two groups, as often as possible, and mixing and matching parents as much as possible.
  4. Also get any available arilpums (pretty much just ‘Aladdin’s Gem’ and ‘Topaz Talisman’ these days), to cross amongst themselves or with arilpum seedlings you produced from the aril/pumila crosses.

Hildebrandt arilpum seedling 11-4-2
(photo: George Hildebrandt)
I think this a project that offers enormous potential in the long term, and is ideal for a small, backyard hybridizer who does not have the resources to grow thousands of seedlings and does not need to worry about immediate commercial success. There is a whole new world waiting to be opened up!

Monday, June 4, 2018

In Praise of Regelias

by Tom Waters

The Regelias are a group of irises native to central Asia, their range extending from near the Caspian Sea to the mountainous regions bordering Tibet. Their nearest relatives are the oncocyclus, which are found further west and south from western Iran to the Mediterranean. Regelias and oncocyclus together comprise the aril irises.

First, a few comments on the name "Regelia". The name honors German botanist Eduard August von Regel, who was director of the botanical garden in St. Petersburg, Russia, late in the nineteenth century. Russian plant explorers of that time were very active in central Asia, seeking out new species and bringing them to attention of European botanists and gardeners. The proper pronunciation of the name is thus Reh-GEH-li-a, although most English speakers have taken to using the pronunciation Reh-JEE-li-a instead. The name should be capitalized, since it comes from a personal name. Spell-checking software likes to change the name to "regalia", which refers to Royal trappings, a blunder that one should be alert for.

Regelias are similar to oncocyclus in having a large cream-colored aril attached to the seed, in going completely dormant in summer, and in preferring arid conditions. Whereas oncocyclus iris have only one bloom per stalk, Regelias usually have two. Regelias have beards on the inside of the standards as well as the falls! Whereas many oncocyclus have large globular blooms with prominent signals, Regelias have more svelte, elongated flowers, often with conspicuous veining. The two groups are interfertile, and there are advanced-generation hybrids between them.

There are eight or more species of Regelias. Historically, the three species grown in European and American gardens were the diploid Iris korolkowii and the tetraploids I. stolonifera and Iris hoogiana. More recently, I. afghanica and I. lineata are also sometimes obtainable. W. R. Dykes regarded Iris hoogiana as the most beautiful of all irises, because of its satiny sheen and elegant form.
Iris hoogiana
Iris stolonifera

'Vera' (RH)






















There are hybrids between the Regelia species, called Regelia hybrids (RH). Two widely known Regelia hybrids are 'Vera' (uncertain parentage, derived from Iris stolonifera, probably crossed with Iris korolkowii), and 'Bronze Beauty Van Tubergen', a stolonifera/hoogiana hybrid registered by the Aril Society International in 2001, but in commerce since the mid-twentieth century.


All Regelias are adaptable to a wider range of climatic conditions than their oncocyclus relatives. Here in northern New Mexico, mine persist better than daffodils, and get the same care. Cold winters present no problem whatsoever, as they are native to continental mountainous regions. Dampness and humidity in summer can cause problems, as the plants are dormant then and susceptible to rot.

Regeliocycli

'Bronze Beauty Van Tubergen' (RH)
Once it was understood that the Regelias and oncocyclus irises could be crossed readily, hybridizers became interested in such hybrids, mostly as way to breed the Regelia adaptability into the often troublesome oncocycli, which are notoriously particular and difficult to grow in many climates. The firm of Van Tubergen produced a number of regeliocyclus hybrids in the early 20th century, many of which indeed proved quite durable and are still enjoyed today. Most of these were produced by crossing Iris korolkowii with oncocyclus species, and showed both Regelia veining and oncyclus dotting and signals. In current usage, the term "regeliocyclus" (RC) refers to a hybrid with both Regelia and oncocyclus ancestry that is predominantly Regelia in appearance. In practice, regeliocycli are aril hybrids with 1/2 Regelia ancestry or more.

'Dardanus' (Van Tubergen, not registered) (RC)
Regeliabreds

When hybridizing interest in arils blossomed in the 1940s and 1950s, the attention was almost exclusively on the oncocyclus. Regelias were thought of as "poor relations" that were not always welcome at the table. The prevailing opinion at the time was that they might be useful in breeding arilbreds that were easier to grow, or to facilitate breeding oncocyclus with bearded irises, but it was the "onco look" that was the holy grail of arilbred breeders, and signs of Regelia ancestry were frowned upon.

'Stars Over Chicago' (H. Danielson, 1973)
Henry Danielson was among the first to produce and promote arilbreds of purely Regelia ancestry, launching a popular series of regeliabreds with 'Genetic Artist' (H. Danielson, 1972). These regeliabreds (RB) were derived mostly from I. stolonifera and I. hoogiana. Rather than the globular oncocyclus look expected of arilbreds at the time, they tended toward elongated, open form, showing off the often dramatic colors of the insides of the standards. Although I. stolonifera itself tends to brownish and muted violet tones, its arilbred descendants often combine gold or yellow color with lavender or electric blue flushes in the center of the falls and standards, with similarly colored beards. These unconventional arilbreds were welcomed enthusiastically by some, but reviled by others as garish departures from the oncocyclus ideal.

In the 21st century, French hybridizer Lawrence Ransom picked up the torch of regeliabred breeding, using the Regelia hybrid 'Vera' to produce the siblings 'Eastern Blush' (2002) and 'Eastern Dusk' (2010). 'Eastern Dusk' then gave the distinctive horned arilbred 'Poisonous' (Ransom, 2010).

'Poisonous' (RB-)
Ransom also produced a delightfully varied series of regeliabred arilbred medians, again using 'Vera' as the Regelia parent, with mixed SDB pollen. These "Vera girls" include the widely grown 'Vera-Marina' (Ransom, 1998) and 'Vera-Ruby' (Ransom, 1996).

A special favorite of mine is an arilbred dwarf regeliabred from I. stolonifera X I. pumila, 'Topaz Talisman' (Jensen, 2015), from long-time Regelia enthusiast Elm Jensen, registered at 10 inches in height.

'Topaz Talisman' (RB)
Regelias and their regeliabred descendants have attracted a devoted following over the years, and have shown their great potential in adding variety, interest, and ease of culture to a collection of arils and arilbreds. Much of their full potential, I believe, is still untested. For decades, they have suffered from unfair comparison with their more popular oncocyclus and oncobred cousins. As more growers and hybridizers move away from the prejudices of the past, the Regelias and regeliabreds may at last come into their own as fascinating and beautiful types of iris in their own right.

Monday, April 23, 2018

Aril Trek 2018!

by Tom Waters

'Point Well Taken' (McAllister, 1998)
at the Pountney garden
On April 6 and 7, aril and arilbred enthusiasts from parts far and wide converged on Las Cruces, New Mexico, to enjoy each other's company and an astonishing array of arilbreds in bloom.

The Aril Trek was a joint undertaking of the Aril Society International and the Mesilla Valley Iris Society, and the arrangements were all excellent and smoothly executed.

A labyrinth patterned after the one at Chartres Cathedral,
in Howard Dash's garden
Friday evening registration was followed an educational session of "aril bingo" and a panel discussion with Dell Perry, Pete McGrath, Rick Tasco, Perry Dyer, and (much to my surprise) yours truly.

'New Vision' (Tasco, 2012)
at the Dash garden
At the Wilson Garden
The heart of the weekend was four garden tours on Saturday: the small but beautifully designed Pountney garden in the residential Tortugas area south of Las Cruces, the expansive Dash garden high in the West Mesa desert, the glorious Wilson Garden north of the city, with row upon abundant row of perfectly grown irises of many types, and the delightfully ornamented Ayres garden, northeast of the city.

The timing seemed perfect for arilbreds, with both halfbreds and quarterbreds apparently near peak bloom, amidst late medians and early TBs. I got to see many old and new cultivars in bloom that I have not grown myself. It was also rewarding to see such fine arilbred performance in gardens with such different growing conditions: desert sand to river-valley clay, cool verdant niches to sites exposed to the full sun, wind, and temperature extremes of southern New Mexico.

'Dubai' (Johnson, 2013)
at the Wilson Garden
Saturday afternoon featured informative judges training led by Dell Perry (who took us through the intricacies of aril and arilbred classification and characteristics with verve and expertise) and Perry Dyer (who applied his experience and keen eye to show how to look at these varied plants in the garden). Yes, I did pass the test at the end!

'Gideon's Lamp' (McGrath, 2004)
at the Wilson Garden
At the Saturday night dinner, keynote speaker Pete McGrath amazed us all with slides of startlingly beautiful pure aril hybrids and tantalizing arilbred seedlings from his continuing effort to bring bolder veining into this group. Pete's talk was also a story of his experience with aril and arilbred irises, full of deep disappointments and unexpected rewards, and always coming back to the personal dimension: the people and relationships that affected him and his work, often in profound ways.

'Mean Mr. Mustard' (McGrath, 2010)
at the Ayres garden
It was a great weekend. I particularly enjoyed renewing old acquaintances in the iris world, and meeting other new friends I had only known on line. If you are an iris lover, but not sure if you can manage the time and expense of attending a national convention, regional and special-interest gatherings such as the Aril Trek are a comfortable and rewarding way to see some beautiful iris gardens and meet many wonderful people!


Monday, January 15, 2018

The Legacy of 'Anacrusis'

by Tom Waters

'Anacrusis' (Mathes, 1992), OGB/ABM
There is a very interesting line of irises created by Harald Mathes of Germany, beginning with 'Anacrusis' (Mathes, 1992). This iris was produced in a very interesting way. Mathes crossed a pure oncocyclus aril hybrid (I. iberica x I. auranitica) with the dwarf bearded species I. suaveolens. Both the aril and the dwarf are diploids (two sets of chromosomes), and wide crosses between diploids ordinarily produce sterile offspring. Mathes, however, used colchicine treatment to convert the seedling to tetraploidy (four sets of chromosomes). Theory predicts that such a tetraploid would be fertile, and indeed it was! (See my earlier blog post, Tetraploid Arils, Anyone?) Tetraploids created in this way can be unstable, reverting to the diploid state or growing poorly or erratically. So to preserve this breeding accomplishment, Mathes used the pollen of the tetraploid seedling on a conventional arilbred, 'Dresden Gold' (Foster, 1975). The result of that cross was 'Anacrusis'.

'Anacrusis' thus carries an unusual complement of genetic material. It has two sets of aril chromosomes, one from the oncocyclus hybrid Mathes had used, and one from 'Dresden Gold'. It has one set of TB chromosomes from 'Dresden Gold', and one set of dwarf suaveolens chromosomes. So it is a "halfbred", with two sets of aril chromosomes and two sets of bearded chromosomes, but with an important difference: one of the bearded sets is from the dwarf species I. suaveolens, rather than the TB ancestry found in most halfbreds. The aril species I. auranitica is also something different, not to be found in the ancestry of most halfbreds.  'Anacrusis', the result of this unprecedented combination of genetic material, is an arilbred median (20 inches in height) with the globular oncocyclus flower form, dark color, and a large black signal.

It is worthwhile to compare this with more typical arilbred medians, which come from crossing halfbreds with standard dwarf bearded irises (SDBs). These get their dwarf ancestry from I. pumila, whereas 'Anacrusis' gets its from I. suaveolens. Conventional arilbred medians are one-quarter aril, but 'Anacrusis' is one-half aril. And finally, conventional arilbred medians are sterile, but 'Anacrusis' is fully fertile, just like tall halfbreds are.

But what to cross it with? The first 'Anacrusis' child to be introduced was 'Invention' (Mathes, 1994), the result of crossing 'Anacrusis' with a sibling. This iris is similar to 'Anacrusis', also 20 inches in height, and with a similar dark color.

'Concerto Grosso' (Mathes, 1998) OGB/ABM
The next avenue to explore was combining 'Anacrusis' with other arilbreds. To this end, Mathes made use of one of his other unusual arilbreds, 'Gelee Royale' (Mathes, 1982). This iris is that most unusual of creatures, a pentaploid (five sets of chromosomes)! Its pod parent was an triploid OGB+ arilbred (2/3 aril complement, with the aril sets coming from I. auranitica and the Regelia I. hoogiana), with 'Dresden Gold' again as the pod parent. Although this is a complicated pedigree, 'Gelee Royale' breeds much like other halfbreds do. Mathes crossed 'Gelee Royale' with its sibling, crossed the result with the 'Anacrusis' sibling, and then finally crossed the result to 'Invention'. The result of this multi-generation hybridizing work was 'Concerto Grosso' (Mathes, 1998). Despite the presence of 'Gelee Royale' (registered at 35.5 inches) in its pedigree, 'Concerto Grosso' is still only 20 inches tall itslef.

'Concerto Grosso' has larger flowers than 'Anacrusis' and 'Invention', and is a rich mahogany color, quite different from most arilbreds. It went on to with the C. G. White medal, the highest award for arilbreds, in 2005.

'Iridescent Orange' (Mathes, 2001), OGB
Mathes had not finished his work with this line, however. 'Anacrusis', 'Invention', and 'Concerto Grosso' are all very dark in color, and Mathes wanted to extend the range of this line to lighter hues. 'Iridescent Orange' (Mathes, 2001) from ('Invention' x 'Gelee Royale') X 'Concerto Grosso' is a lovely orange self with a dark signal. At 23 inches in height, it is just above the limit for the arilbred median category, as defined in the Checklist of Arilbred Dwarfs and Medians. Its sibling, 'Suprassing Yellow' (Mathes, 2001) is a yellow rendition of the theme, also 23 inches.

The line is carried on in 'Glittering Garnets' (Donald Eaves by Elm Jensen, 2010) is from 'Anacrusis' crossed with the arilbred 'Desert Plum'. This is 22 inches in height, at the upper limit of the arilbred median category.

These irises have also been used in crosses outside the halfbred fertile family to which they belong. 'Dotted Sunsuit' (Mathes, 2001), is an OGB+ triploid from a yellow 'Anacrusis' sib crossed with an oncogelia seedling. 'Chain Reaction' (Tasco, 2007) comes from an SDB seedling x 'Concerto Grosso', a small arilbred median at 13 inches. 'Arcanum' (Jensen, 2013) is from 'Anacrusis' x I. pumila 'Crouching Tiger', a diminutive arilbred dwarf at only 6 inches in height!

There is still potential to be tapped from the 'Anacrusis' legacy. Its value in breeding fertile arilbred medians should be noticed and pursued. The most promising way to go about this is to cross 'Anacrusis' or any of its descendants with the smallest halfbreds available. 'Peresh' (Whitely, 2001), at 15 inches, and its siblings 'Eglon', 'Kedesh', and 'Tekoah' come to mind.

The creative, unprecedented cross that produced 'Anacrusis' is a true hybridizing success story, not only producing an interesting, attractive iris, but opening up new paths for the hybridizer and for others who were to follow.




Monday, October 16, 2017

An Arilbred Iris Sampler

Tom Waters

'Oyez' (White, 1938)
I was recently contacted by an iris lover who had read my blog post on the classification of arilbred irises and wanted to sample each of the categories. She asked if I had any recommendations. I gave a few off the top of my head, and she suggested that this might make a good blog topic. I agree! So here is my arilbred sampler, for those who want to try the full range of different types of arilbreds.

I've restricted the irises mentioned here to those that are readily available. The commercial gardens that have the best selection of arilbreds are Superstition Iris Gardens, Mid-America Garden, and Blue J Iris. In addition, the Aril Society International has an annual plant sale with many more arilbreds than these commercial growers offer. The offerings vary each year depending on what members contribute, but many varieties are offered nearly every year. If an iris was available from one of the three large commercial growers in their most recent catalog, or in the ASI plant sales for the last two years, I deemed it readily available. Some other commercial growers list a small number of arilbreds, and if arilbreds are grown in your area, the local iris society plant sales can also be a good source of plants.

I want to be clear that this listing is indeed intended as a "sampler". The idea is to cover as many different kinds as possible, to showcase the diversity of arilbreds. It is not an attempt to identify the "best arilbreds" by some objective criteria, nor even a list of "Tom's favorites". The idea is that if you are new to arilbreds, and plant a varied assortment, you can discover your own preferences and what grows well for you.

The ASI recognizes nine different categories of arilbred, based on the type and amount of aril content the iris has. There are two types of pure arils: oncocyclus and Regelia. An arilbred can have either or both of these types of aril in its background. Hence we have oncobreds (OB), Regeliabreds (RB), and oncogeliabreds (OGB, mixed onco and Regelia content). Furthermore, arilbreds can be half aril, more than half aril, or less than half aril. Those with more than half aril content are indicated with a "+" sign; those with less than half aril with a "-" sign. No sign is used for those that are half aril. Thus RB+ means an arilbred whose type of aril content is Regelia only, and whose amount of aril content is more than half.


It turns out that these nine categories actually allow for more distinctions than a practical gardener need attend to. There are very few OB arilbreds, and they are indistinguishable from most OGB types. The reason is that most of today's arilbreds are descended from pioneering work by C. G. White in the 1940s, and White did not keep track of parentages during this time. So his arilbreds, although drawing heavily on oncocyclus arils and selected to resemble oncocyclus as much as possible, are classified as OGB because of the possibility of some Regelia ancestry. So I don't separate OBs from OGBs here.

I do, however, separate out the arilbred medians and arilbred dwarfs from the tall arilbreds. These smaller arilbreds, typically with dwarf or median bearded irises in their parentage, fill a very different role in the garden and provide much of the diversity to be encountered in the arilbred tribe.

So much for the preliminaries. Onward to the plants!


OGB and OB (Onco-type halfbreds)

These are what most of us think of when we think of arilbreds. Most are tall types, with TB and oncocyclus ancestry dominating. Blooms are typically large and globular, often with strong aril markings, such as signal spots or veining.

'Refiner's Fire' (McGrath, 2007)
Large bold signals are actually a rather recent development in this group, with 'Energizer' (Shockey, 1996) being a striking breakthrough in its time. More recently, 'Noble Warrior' (Tasco, 2006) combines a dramatic signal with a bit of veining. 'Refiner's Fire' (McGrath, 2007) gives a striking dark signal on a deeply colored red-toned flower.

'Sand Dancer' (Tasco, 2010) gives us a whole different look, with falls covered in peppery stippling. Yet another completely different look is to be found in 'Navajo Velvet' (McAllister, 2009), with falls of lustrous, satiny mahogany red.

The historic arilbred 'Oyez' (White, 1938) has seduced many iris growers into the arilbred world with its dramatic precise veining. Strangely, it has been difficult to reproduce this striking pattern in modern arilbreds. ('Oyez' is a sterile diploid.) An early success was 'Jonnye's Magic' (Rich, 1992). For a true modern advancement in veined arilbreds, seek out 'Alakazam' (Tasco, 2013).

'Aladdin's Gem' (Thoolen, 2002)
There are few arilbred medians and arilbred dwarfs in this category. Very notable is 'Aladdin's Gem' (Thoolen, 2002), a fertile aril-pumila hybrid about the size of a small SDB (standard dwarf bearded)!

OGB- and OB- (Onco-type quarterbreds)

Originally, these categories were dominated by tall arilbreds that came from crossing halfbreds with TBs. At a time when halbreds were uncommon and difficult to grow, the quarterbreds were the most accessible arilbreds for most gardeners. The historic iris 'Elmohr' (Loomis-Long, 1942) was a fine example of the type, the only arilbred to win the Dykes Medal. It does not show the strong aril patterning we now expect in arilbreds, without signal or veining to speak of. To understand its allure, we must think back to a time when TBs were troubled with narrow petals and often droopy form. 'Elmohr' stood out with dramatically large blooms, wide falls, and a bit of globular onco form to bring it all together.

Some other tall quarterbreds for the sampler are 'Dune' (Hager, 1981) and the very recent 'Heart of Hearts' (Black, 2015). At their best, this type of iris can bring some of the modern TB form and bud count into irises that still evoke their aril ancestry.

'Desert Snow' (Black, 2013)
Since the 1970s, however, arilbred medians have increasingly dominated these categories. Crossing an
OGB halfbred with an SDB produces an OGB- arilbred median. In the garden, these occupy a niche similar to the IBs (intermediate beardeds), but with some aril traits to give them that special something extra. The charm and adaptability of this type of arilbred has given them a great appeal among arilbred growers and median fanciers alike. 'Brash and Bold' (Black, 2009) and 'Desert Snow' (Black, 2013) are fine examplars of what this category has to offer.

A somewhat different approach is found in 'Persian Sapphire' (Baumunk, 2005), a child of 'Aladdin's Gem' that has more Iris pumila in its makeup than TB.

There are some arilbred dwarfs in this category that are worthy of attention. 'Loudmouth' (Rich, 1970) is a perennial favorite, SDB-sized with raucous signal and veining and globular form. 'Tiny Pirate' (Rich, 1990) is the most diminutive arilbred I have grown; it would be small even among MDBs (miniature dwarf beardeds), but struts onco form, a signal, and a whisper of veining.

RB (Regelia-type halfbreds)

Enthusiasm for large, globular oncocylcus irises with their dramatic signals and stippling was a driving force in early arilbred breeding. The Regelias took a back seat in the minds of many growers and breeders; sometimes they were thought of as providing nothing but some ruggedness and climate adaptability. But some have always appreciated the Regelias for their svelte elegance, sometimes striking veining, satiny texture, or blended colors.

An early Regeliabred that helped draw attention to the potential of this type of iris was 'Genetic Artist' (H. Danielson, 1972). This shows a classic color pattern derived from the Regelia Iris stolonifera: a yellowish rim around a blue or violet center. 'Afrosiab' (Volfovich-Moler, 2001) shows a touch of ruffling from its TB parent, 'Mary Frances' (Gaulter, 1973). The French hybridizer Lawrence Ransom worked extensively with Regeliabreds, with his 'Eastern Blush' (Ransom, 2002) being much used in his breeding program.

RB- (Regelia-type quarterbreds)

Ransom's work takes center stage in this category. Among the tall RB- are the horned 'Sandthorn' (Ransom, 2011), and his "Pashtun" series, for example 'Pashtun Princess' (Ransom, 2011).

Ransom also produced a delightfully varied series of RB- arilbred medians, the "Vera girls", from crossing the Regelia 'Vera' (Van Tubergen, not registered) with SDBs.of which 'Vera-Marina'  (Ransom, 1998) is one example.

OGB+ (Onco-type "three-quarter"-breds)

This group has become sadly scarce in commerce, with seldom any new ones registered and introduced. This is perhaps because few arilbred hybridizers today grow the oncocyclus species and hybrids needed to produce this type of arilbred, which usually comes from crossing an OGB arilbred with a pure oncocyclus. The few that are readily available are not always representative of the best this type of breeding has to offer. 'Tul Kerem' (H. Danielson, 1974) is interesting but I find its combination of oncocylcus and Regelia traits ends up not doing justice to either. 'Masada's Glory' (Whitely, 2002) is a better exemplar of what this category has to offer. Although they do not meet my availability criteria, keep a look out from 'Jeweled Veil' (Rich, 1978) or 'Dotted Sunsuit' (Mathes, 2001). These show the oncocyclus features of their ancestry to best advantage.

RB+ (Regelia-type "three-quarter"-breds)

'Turkish Topaz' (Austin, 1962)
A couple oddities round out the sampler. 'Turkish Topaz' (Austin, 1962) was registered as a pure Regelia hybrid, but its parentage is somewhat ambiguous and it looks and grows like an RB+. The flowers are yellow with much brown streaking and blotching. 'White Arts' (L. Danielson, 1986) has only Iris hoogiana in its Regelia ancestry. This species shows nothing that we might recognize as distinctively aril in its coloration. If you walked by 'White Arts' in a garden, you could be forgiven for thinking it was a white IB or perhaps an antique diploid TB.

If you want to sample the world of arilbreds, be sure to try several different types. The variety available in plant size, color pattern, and form is truly remarkable!